esinevad taimedes, sisaldavad spetsiifilist WRKY domeeni, mis seostub märklaudgeenide DNA W-box (T/A)TGAC(T/A) järjestusega cis-elemendiga, WRKY-domain containing proteins, bind to the DNA W-box sequence cis-element. Required for the polarization of the zygote, stress responses and plant development.