paljudest erinevatest alaühikutest koosnevad roteeruvad molekulaarsed kompleksid mitokondrite sisemembraanis ja kloroplastide tülakoidide membraanides, mis teostavad ATP sünteesi elektrontransportahelates tekkinud prootonite liikumapaneva jõu arvel, Rotary molecular complexes located in the inner membranes of mitochondria and thylacoids. Use a proton gradient to drive ATP synthesis by allowing the passive flux of protons across the membrane down their electrochemical gradient and using the energy released by the transport reaction to release newly formed ATP from the active site. Structurally similar to V-type ATPases, contain a FO (intracellular) and F1 (extracellular) area in mitochondria and CFO and CF1 area in chloroplasts, respectively.